Animalia > Chordata > Aves > Passeriformes > Muscicapidae > Ficedula > Ficedula hypoleucaFicedula hypoleuca (European Pied Flycatcher)Synonyms: Motacilla hypoleuca (homotypic); Muscicapa atricapilla; Muscicapa hypoleuca The European pied flycatcher (Ficedula hypoleuca) is a small passerine bird in the Old World flycatcher family. One of the four species of Western Palearctic black-and-white flycatchers, it hybridizes to a limited extent with the collared flycatcher. It breeds in most of Europe and western Asia. It is migratory, wintering mainly in western Africa. It usually builds its nests in holes on oak trees.This species practices polygyny, usually bigamy, with the male travelling large distances to acquire a second mate. The male will mate with the secondary female and then return to the primary female in order to help with aspects of child rearing, such as feeding. |
Uniqueness Scale: Similiar (0) Unique (100) Uniqueness & Vulnerability Scale: Similiar & Secure (0) Unique & Vulnerable (100) ED Score: 4.40631 EDGE Score: 1.68757 |
Clutch Size [8] | 6 | Clutches / Year [9] | 1 | Fledging [5] | 16 days | Incubation [7] | 14 days | Mating Display [3] | Ground and non-acrobatic aerial display | Mating System [3] | Monogamy (mostly) | Maximum Longevity [6] | 15 years | Migration [1] | Intercontinental | Water Biome [1] | Coastal | Wing Span [9] | 9 inches (.22 m) |  | Adult Weight [2] | 13 grams | Birth Weight [3] | 1.7 grams |  | Diet [4] | Carnivore (Invertebrates) | Diet - Invertibrates [4] | 100 % | Forages - Canopy [4] | 33 % | Forages - Mid-High [4] | 33 % | Forages - Understory [4] | 33 % |  | Female Maturity [6] | 1 year | Male Maturity [6] | 1 year |
|
Name |
Countries |
Ecozone |
Biome |
Species |
Report |
Climate |
Land Use |
Alps conifer and mixed forests |
Italy, France, Switzerland, Slovenia |
Palearctic |
Temperate Coniferous Forests |
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Atlantic Equatorial coastal forests |
Cameroon, Congo, Democratic Republic of the Congo, Equatorial Guinea, Gabon |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
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Atlantic mixed forests |
France, Belgium, Netherlands, Germany, Denmark |
Palearctic |
Temperate Broadleaf and Mixed Forests |
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Baltic mixed forests |
Germany, Denmark, Sweden, Poland |
Palearctic |
Temperate Broadleaf and Mixed Forests |
|
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|
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Caledon conifer forests |
United Kingdom |
Palearctic |
Temperate Coniferous Forests |
|
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Cameroonian Highlands forests |
Cameroon, Nigeria |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
|
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Cantabrian mixed forests |
Spain, Portugal |
Palearctic |
Temperate Broadleaf and Mixed Forests |
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Carpathian montane forests |
Czech Republic, Poland, Romania, Slovakia, Ukraine |
Palearctic |
Temperate Coniferous Forests |
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Celtic broadleaf forests |
Ireland, United Kingdom |
Palearctic |
Temperate Broadleaf and Mixed Forests |
|
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Central European mixed forests |
Austria, Belarus, Czech Republic, Germany, Lithuania, Moldovia, Poland, Romania, Russia, Ukraine |
Palearctic |
Temperate Broadleaf and Mixed Forests |
|
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|
|
Cross-Niger transition forests |
Nigeria |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
|
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Cross-Sanaga-Bioko coastal forests |
Cameroon, Equatorial Guinea, Nigeria |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
|
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|
|
East European forest steppe |
Bulgaria, Moldova, Romania, Russia, Ukraine |
Palearctic |
Temperate Broadleaf and Mixed Forests |
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|
|
East Sudanian savanna |
Chad, Central African Republic, Sudan, Ethiopia, Uganda |
Afrotropic |
Tropical and Subtropical Grasslands, Savannas, and Shrublands |
|
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|
|
Eastern Guinean forests |
Ghana, Côte d'Ivoire, Benin, Togo |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
|
|
|
|
English Lowlands beech forests |
United Kingdom |
Palearctic |
Temperate Broadleaf and Mixed Forests |
|
|
|
|
Guinean forest-savanna mosaic |
Nigeria, Benin, Togo, Ghana, Côte d'Ivoire, Guinea, Sierra Leone, Guinea-Bissau, Gambia, Senegal |
Afrotropic |
Tropical and Subtropical Grasslands, Savannas, and Shrublands |
|
|
|
|
Guinean montane forests |
Guinea, Côte d'Ivoire, Liberia, Sierra Leone |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
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|
|
Iberian conifer forests |
Spain |
Palearctic |
Mediterranean Forests, Woodlands, and Scrub |
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Iberian sclerophyllous and semi-deciduous forests |
Spain, Portugal |
Palearctic |
Mediterranean Forests, Woodlands, and Scrub |
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|
Mandara Plateau mosaic |
Cameroon, Nigeria |
Afrotropic |
Tropical and Subtropical Grasslands, Savannas, and Shrublands |
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|
Mediterranean acacia-argania dry woodlands and succulent thickets |
Morocco, Spain |
Palearctic |
Mediterranean Forests, Woodlands, and Scrub |
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|
|
Mediterranean conifer and mixed forests |
Algeria, Morocco, Tunisia |
Palearctic |
Temperate Coniferous Forests |
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|
Mediterranean High Atlas juniper steppe |
Morocco |
Palearctic |
Montane Grasslands and Shrublands |
|
|
|
|
Mediterranean woodlands and forests |
Morocco, Algeria, Tunisia |
Palearctic |
Mediterranean Forests, Woodlands, and Scrub |
|
|
|
|
Mount Cameroon and Bioko montane forests |
Cameroon, Equatorial Guinea |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
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|
|
Niger Delta swamp forests |
Nigeria |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
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|
|
Nigerian lowland forests |
Nigeria, Benin |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
|
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|
|
North Atlantic moist mixed forests |
Ireland, United Kingdom |
Palearctic |
Temperate Broadleaf and Mixed Forests |
|
|
|
|
Northeastern Congolian lowland forests |
Cameroon, Central African Republic, Gabon, Republic of the Congo |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
|
|
|
|
Northeastern Spain and Southern France Mediterranean forests |
France, Spain |
Palearctic |
Mediterranean Forests, Woodlands, and Scrub |
|
|
|
|
Northern Congolian forest-savanna mosaic |
Democratic Republic of the Congo, Sudan, Central African Republic, Cameroon |
Afrotropic |
Tropical and Subtropical Grasslands, Savannas, and Shrublands |
|
|
|
|
Northwest Iberian montane forests |
Portugal, Spain |
Palearctic |
Mediterranean Forests, Woodlands, and Scrub |
|
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|
|
Northwestern Congolian lowland forests |
Cameroon, Central African Republic, Gabon, Democratic Republic of the Congo |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
|
|
|
|
Pannonian mixed forests |
Austria, Bosnia & Herzegovina, Croatia, Czech Republic, Hungary, Romania, Serbia, Slovakia, Slovenia, Ukraine |
Palearctic |
Temperate Broadleaf and Mixed Forests |
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|
Pontic steppe |
Russia, Kazakhstan, Ukraine |
Palearctic |
Temperate Grasslands, Savannas, and Shrublands |
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|
Red Sea coastal desert |
Egypt, Sudan |
Palearctic |
Deserts and Xeric Shrublands |
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Sarmatic mixed forests |
Belarus, Estonia, Finland, Latvia, Lithuania, Norway, Russia, Sweden |
Palearctic |
Temperate Broadleaf and Mixed Forests |
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Sayan Alpine meadows and tundra |
Mongolia, Russia |
Palearctic |
Montane Grasslands and Shrublands |
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Scandinavian and Russian taiga |
Norway, Sweden, Finland, Russia |
Palearctic |
Boreal Forests/Taiga |
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Scandinavian coastal conifer forests |
Norway |
Palearctic |
Temperate Coniferous Forests |
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Scandinavian Montane Birch forest and grasslands |
Norway, Sweden, Finland |
Palearctic |
Tundra |
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Southeastern Iberian shrubs and woodlands |
Spain |
Palearctic |
Mediterranean Forests, Woodlands, and Scrub |
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Southwest Iberian Mediterranean sclerophyllous and mixed forests |
Portugal, Spain, France, Italy, Morocco |
Palearctic |
Mediterranean Forests, Woodlands, and Scrub |
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|
|
West Sudanian savanna |
Senegal, Guinea, Mali, Côte d'Ivoire, Burkina Faso, Ghana, Togo, Benin, Niger, Nigeria |
Afrotropic |
Tropical and Subtropical Grasslands, Savannas, and Shrublands |
|
|
|
|
Western Congolian forest-savanna mosaic |
Gabon, Congo, Democratic Rep of the Congo, Angola |
Afrotropic |
Tropical and Subtropical Grasslands, Savannas, and Shrublands |
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|
|
Western Congolian swamp forests |
Democratic Republic of the Congo, Congo |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
|
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|
|
Western European broadleaf forests |
Austria, Czech Republic, France, Germany |
Palearctic |
Temperate Broadleaf and Mixed Forests |
|
|
|
|
Western Guinean lowland forests |
Guinea, Côte d'Ivoire, Liberia, Sierra Leone |
Afrotropic |
Tropical and Subtropical Moist Broadleaf Forests |
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|
Name |
IUCN Category |
Area acres |
Location |
Species |
Website |
Climate |
Land Use |
Alsó-Tisza-völgy |
|
88306 |
Hungary |
|
|
|
|
Alte Valli Stura e Maira |
|
103806 |
Italy |
|
|
|
|
Ammergebirge mit Kienberg und Schwarzenberg sowie Falkenstein |
|
74415 |
Germany |
|
|
|
|
Astrakhanskiy Biosphere Reserve Zapovednik |
Ia |
167827 |
Astrakhan, Russia |
|
|
|
|
Belovezhskaya Pushcha Biosphere Reserve National Park |
II |
218515 |
Belarus |
|
|
|
|
Berezinskiy Zapovednik State Nature Reserve |
Ia |
209113 |
Belarus |
|
|
|
|
CAMPIÑAS DE SEVILLA |
|
88303 |
Spain |
|
|
|
|
Camargue |
|
546256 |
France |
|
|
|
|
Chausey |
|
203679 |
France |
|
|
|
|
Chernye Zemli Biosphere Reserve Zapovednik |
IV |
621951 |
Russia |
|
|
|
|
Corbett National Park |
II |
141853 |
Uttaranchal, India |
|
|
|
|
DELTA ACHELOOU, LIMNOTHALASSA MESOLONGIOU - AITOLIKOU KAI EKVOLES EVINOU, NISOI ECHINADES, NISOS PET |
|
109185 |
Greece |
|
|
|
|
DELTA ACHELOOU, LIMNOTHALASSA MESOLONGIOU - AITOLIKOU, EKVOLES EVINOU, NISOI ECHINADES, NISOS PETALA |
|
87747 |
Greece |
|
|
|
|
DELTA AXIOU - LOUDIA - ALIAKMONA - ALYKI KITROUS |
|
73260 |
Greece |
|
|
|
|
DELTA AXIOU - LOUDIA - ALIAKMONA - EVRYTERI PERIOCHI - AXIOUPOLI |
|
83216 |
Greece |
|
|
|
|
DESFILADEROS DEL RÍO MARTÍN |
|
110851 |
Spain |
|
|
|
|
DOÑANA |
|
281453 |
Spain |
|
|
|
|
Danube Delta Biosphere Reserve |
|
1433211 |
Romania |
|
|
|
|
Darvinsky Zapovednik |
Ia |
278421 |
Yaroslavl, Vologda, Russia |
|
|
|
|
Douro Internacional e Vale do Águeda |
|
125502 |
Portugal |
|
|
|
|
Eastern Beskid National Park |
II |
68484 |
Poland |
|
|
|
|
Estuário do Tejo |
|
110634 |
Portugal |
|
|
|
|
Felsõ-kiskunsági szikes puszták és turjánvidék |
|
103542 |
Hungary |
|
|
|
|
Flusslandschaft Elbe Biosphere Reserve |
|
925242 |
Germany |
|
|
|
|
Hortobágy National Park |
II |
140004 |
Hungary |
|
|
|
|
Itäisen Suomenlahden saaristo ja vedet |
|
236302 |
Finland |
|
|
|
|
KORYFES OROUS GRAMMOS |
|
84898 |
Greece |
|
|
|
|
Kiskunsági szikes tavak és az õrjegi turjánvidék |
|
88335 |
Hungary |
|
|
|
|
Krivoklatsko Protected Landscape Area |
V |
155267 |
Czech Republic |
|
|
|
|
Krkonose National Park |
V |
95995 |
Czech Republic |
|
|
|
|
LA SERENA Y SIERRAS PERIFÉRICAS |
|
379806 |
Spain |
|
|
|
|
LEFKA ORI KAI PARAKTIA ZONI |
|
131864 |
Greece |
|
|
|
|
LOS VALLES |
|
79029 |
Spain |
|
|
|
|
Lagorai |
|
114140 |
Italy |
|
|
|
|
Lake District High Fells |
|
66717 |
England, United Kingdom |
|
|
|
|
Lake Torne Area Biosphere Reserve |
|
238457 |
Sweden |
|
|
|
|
Laplandskiy Biosphere Reserve |
|
757114 |
Russia |
|
|
|
|
Les Cévennes |
|
227446 |
France |
|
|
|
|
MATARRAÑA - AIGUABARREIX |
|
90747 |
Spain |
|
|
|
|
MONTES DE ZUERA, CASTEJÓN DE VALDEJASA Y EL CASTELLAR |
|
63037 |
Spain |
|
|
|
|
MONTES OBARENES |
|
106406 |
Spain |
|
|
|
|
MONTES UNIVERSALES - SIERRA DEL TREMEDAL |
|
79316 |
Spain |
|
|
|
|
Marahoue National Park |
II |
293884 |
Côte d'Ivoire |
|
|
|
|
Merenkurkun saaristo |
|
316695 |
Finland |
|
|
|
|
Montesinho/Nogueira |
|
266900 |
Portugal |
|
|
|
|
Moor House – Upper Teesdale |
|
95867 |
England, United Kingdom |
|
|
|
|
Mordovsky Zapovednik |
Ia |
79439 |
Mordovia, Russia |
|
|
|
|
Morecambe Bay |
|
151985 |
England, United Kingdom |
|
|
|
|
NOTIO DASIKO SYMPLEGMA EVROU |
|
72341 |
Greece |
|
|
|
|
NOTIODYTIKI CHERSONISOS, APOLITHOMENO DASOS LESVOU |
|
71214 |
Greece |
|
|
|
|
North Pennine Moors |
|
254789 |
England, United Kingdom |
|
|
|
|
North York Moors |
|
108930 |
England, United Kingdom |
|
|
|
|
OREINOS EVROS - KOILADA DEREIOU |
|
120853 |
Greece |
|
|
|
|
ORI ANATOLIKIS LAKONIAS |
|
92829 |
Greece |
|
|
|
|
OROS OTHRYS, VOUNA GKOURAS KAI FARANGI PALAIOKERASIAS |
|
76799 |
Greece |
|
|
|
|
OROS PARNASSOS |
|
84965 |
Greece |
|
|
|
|
Okskiy Biosphere Reserve |
|
190748 |
Russia |
|
|
|
|
PARAPOTAMIO DASOS VOREIOU EVROU KAI ARDA |
|
64079 |
Greece |
|
|
|
|
POSETS - MALADETA |
|
82205 |
Spain |
|
|
|
|
Parc National de Tai National Park |
II |
1085310 |
Côte d'Ivoire |
|
|
|
|
Parc National de la Comoe National Park |
II |
2902593 |
Côte d'Ivoire |
|
|
|
|
Parc National du Niokolo-Koba National Park |
II |
2046878 |
Senegal |
|
|
|
|
Parco Del Cilento e Vallo de Diano National Park |
|
551095 |
Italy |
|
|
|
|
Parco Regionale Orobie Bergamasche |
|
121015 |
Italy |
|
|
|
|
Parco delle Madonie |
|
100967 |
Italy |
|
|
|
|
Pechoro-Ilychskiy Biosphere Reserve |
|
3098091 |
Russia |
|
|
|
|
Pflaelzerwald Nature Park |
V |
438818 |
Germany |
|
|
|
|
Puszcza Kampinoska Biosphere Reserve |
|
188373 |
Poland |
|
|
|
|
Reserve de la Biosphere des Vosges du Nord Biosphere Reserve |
|
301469 |
France |
|
|
|
|
Rios Sabor e Maçãs |
|
125253 |
Portugal |
|
|
|
|
Risaie della Lomellina |
|
76457 |
Italy |
|
|
|
|
RÍO GUADALOPE - MAESTRAZGO |
|
133800 |
Spain |
|
|
|
|
RÍO HUERVA Y LAS PLANAS |
|
74804 |
Spain |
|
|
|
|
SABINARES DEL ARLANZA |
|
92427 |
Spain |
|
|
|
|
SIERRA DE ALCUBIERRE |
|
103846 |
Spain |
|
|
|
|
SIERRA DE GRAZALEMA |
|
131980 |
Spain |
|
|
|
|
SIERRA DE LA DEMANDA |
|
125797 |
Spain |
|
|
|
|
SIERRA NEVADA |
|
424554 |
Spain |
|
|
|
|
SIERRA SUBBETICA |
|
79212 |
Spain |
|
|
|
|
SIERRAS DE CAZORLA, SEGURA Y LAS VILLAS |
|
519222 |
Spain |
|
|
|
|
SIERRAS DE SANTO DOMINGO Y CABALLERA Y RÍO ONSELLA |
|
88167 |
Spain |
|
|
|
|
Saaristomeri |
|
147873 |
Finland |
|
|
|
|
Schorfheide-Chorin Biosphere Reserve |
|
319164 |
Germany |
|
|
|
|
Serra do Gerês |
|
156759 |
Portugal |
|
|
|
|
Severn Estuary/ Môr Hafren |
|
182155 |
England/Wales, United Kingdom |
|
|
|
|
Solway Firth |
|
107829 |
England/Scotland, United Kingdom |
|
|
|
|
South Pennine Moors |
|
160577 |
England, United Kingdom |
|
|
|
|
Spreewald Biosphere Reserve |
|
117355 |
Germany |
|
|
|
|
Stora Sjofallet National Park |
II |
314943 |
Sweden |
|
|
|
|
Sumava Biosphere Reserve |
|
412955 |
Czech Republic |
|
|
|
|
Takamanda Forest Reserve National Park |
II |
167041 |
Cameroon |
|
|
|
|
Tejo Internacional, Erges e Pônsul |
|
63692 |
Portugal |
|
|
|
|
Trebon Basin Protected Landscape Area |
V |
173393 |
Czech Republic |
|
|
|
|
Tsentral'no-Lesnoy Biosphere Reserve |
Ia |
62630 |
Russia |
|
|
|
|
Tsentral'no-Sibirskiy Biosphere Reserve |
Ia |
2521510 |
Krasnoyarsk, Russia |
|
|
|
|
UK-PUISTO-SOMPIO-KEMIHAARA |
|
765461 |
Finland |
|
|
|
|
Ubsunurskaya Kotlovina (Ubsunur Depression) Zapovednik |
Ia |
798640 |
Tuva, Russia |
|
|
|
|
VIÑAMALA |
|
62053 |
Spain |
|
|
|
|
Voronezhskiy Biosphere Reserve |
|
95835 |
Russia |
|
|
|
|
Waddensea of Lower Saxony Biosphere Reserve |
|
539522 |
Germany |
|
|
|
|
Name |
Location |
IBA Criteria |
Website |
Climate |
Land Use |
Brycalavičskaja Pušča |
Belarus |
B2, B3 |
|
|
|
Buzulukski forest |
Russia (European) |
A1, B2, B3 |
|
|
|
Lemmenjoki-Hammastunturi-Pulju |
Finland |
A3, A4i, A4iii, B1i, B2, B3, C2, C3, C4, C6 |
|
|
|
Oulanka-Sukerijärvi |
Finland |
A3, B2, B3, C2, C6 |
|
|
|
Pallas and Ylläs fjelds |
Finland |
A3, B2, B3, C2, C6 |
|
|
|
Pomokaira-Koitelaiskaira |
Finland |
A4i, B1i, B2, B3, C2, C3, C6 |
|
|
|
|
Name |
Location |
Endemic |
Species |
Website |
Caucasus |
Armenia, Azerbaijan, Georgia, Iran, Russia, Turkey |
No |
|
|
Guinean Forests of West Africa |
Benin, Côte d'Ivoire, Equatorial Guinea, Ghana, Guinea, Liberia, Nigeria, São Tomé and Príncipe, Sierra Leone, Togo |
No |
|
|
Mediterranean Basin |
Algeria, Egypt, France, Greece, Israel, Jordan, Lebanon, Libya, Portugal, Spain, Syria, Tunisia, Turkey |
No |
|
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Attributes / relations provided by ♦ 1Myers, P., R. Espinosa, C. S. Parr, T. Jones, G. S. Hammond, and T. A. Dewey. 2006. The Animal Diversity Web (online). Accessed February 01, 2010 at animaldiversity.org♦ 2Storchová, Lenka; Hořák, David (2018), Data from: Life-history characteristics of European birds, Dryad, Dataset, https://doi.org/10.5061/dryad.n6k3n ♦ 3Terje Lislevand, Jordi Figuerola, and Tamás Székely. 2007. Avian body sizes in relation to fecundity, mating system, display behavior, and resource sharing. Ecology 88:1605 ♦ 4Hamish Wilman, Jonathan Belmaker, Jennifer Simpson, Carolina de la Rosa, Marcelo M. Rivadeneira, and Walter Jetz. 2014. EltonTraits 1.0: Species-level foraging attributes of the world's birds and mammals. Ecology 95:2027 ♦ 5Nathan P. Myhrvold, Elita Baldridge, Benjamin Chan, Dhileep Sivam, Daniel L. Freeman, and S. K. Morgan Ernest. 2015. An amniote life-history database to perform comparative analyses with birds, mammals, and reptiles. Ecology 96:3109 ♦ 6de Magalhaes, J. P., and Costa, J. (2009) A database of vertebrate longevity records and their relation to other life-history traits. Journal of Evolutionary Biology 22(8):1770-1774 ♦ 7Intrinsic aging-related mortality in birds, Robert E. Ricklefs, JOURNAL OF AVIAN BIOLOGY 31: 103–111. Copenhagen 2000 ♦ 8Jetz W, Sekercioglu CH, Böhning-Gaese K (2008) The Worldwide Variation in Avian Clutch Size across Species and Space PLoS Biol 6(12): e303. doi:10.1371/journal.pbio.0060303 ♦ 9British Trust for Ornithology♦ 10Ecology of Commanster♦ 11Frugivorous diet of autumn migrant Pied Flycatchers Ficedula hypoleuca: a review and new data, A. Hernández, Butll. GCA 16: 53-60, 1999 ♦ 12Farwig, Nina; Schabo, Dana G.; Albrecht, Jörg (2017), Data from: Trait-associated loss of frugivores in fragmented forest does not affect seed removal rates, Dryad, Dataset, https://doi.org/10.5061/dryad.2784g ♦ 13STATUS, DISTRIBUTION, AND DIET OF ELEONORAS FALCON (FALCO ELEONORAE) IN THE CANARY ISLANDS, Leandro De León, Beneharo Rodríguez, Aurelio Martín, Manuel Nogales, Jesús Alonso, and Carlos Izquierdo, Journal of Raptor Research 41(4):331-336 2007 ♦ 14Gibson, D. I., Bray, R. A., & Harris, E. A. (Compilers) (2005). Host-Parasite Database of the Natural History Museum, London ♦ 15International Flea DatabaseEcoregions provided by World Wide Fund For Nature (WWF). WildFinder: Online database of species distributions, ver. 01.06 Wildfinder Database |
Species taxanomy provided by GBIF Secretariat (2022). GBIF Backbone Taxonomy. Checklist dataset https://doi.org/10.15468/39omei accessed via GBIF.org on 2023-06-13; License: CC BY 4.0
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