Attributes / relations provided by
♦ ¹Roosting Behavior of Colonial and Solitary Flying Foxes in American Samoa (Chiroptera: Pteropodidae), Anne P. Brooke, Christopher Solek, Ailao Tualaulelei, Biotropica 32(2): 338-350, 2000
♦ ²Pteropus hypomelanus, Deborah P. Jones and Thomas H. Kunz, MAMMALIAN SPECIES No. 639, pp. 1–6 (2000)
♦ ³Pteropus tonganus, Carrie A. Miller and Don E. Wilson, MAMMALIAN SPECIES No. 552, pp. 1-6 (1997)