Attributes / relations provided by
♦ ¹del Hoyo, J., Elliott, A., Sargatal, J., Christie, D.A. & de Juana, E. (eds.). Handbook of the Birds of the World Alive. Lynx Edicions, Barcelona.
♦ ²Fruit-frugivore interactions in a Malagasy littoral forest: a community-wide approach of seed dispersal, An Bollen, UNIVERSITEIT ANTWERPEN 2003