Attributes / relations provided by
♦ ¹Tropical Fruit-Eating Birds and Their Food Plants: A Survey of a Costa Rican Lower Montane Forest, Nathaniel T. Wheelwright, William A. Haber, K. Greg Murray, Carlos Guindon, Biotropica Vol. 16, No. 3 (Sep., 1984), pp. 173-192
♦ ²Food niche overlap among neotropical frugivorous bats in Costa Rica, Jorge E. Lopez & Christopher Vaughan, Rev. Biol. Trop. (Int. J. Trop. Biol. ISSN-0034-7744) Vol. 55 (1): 301-313, March 2007
♦ ³"Fig-eating by vertebrate frugivores: a global review", MIKE SHANAHAN, SAMSON SO, STEPHEN G. COMPTON and RICHARD CORLETT, Biol. Rev. (2001), 76, pp. 529–572