Attributes / relations provided by
♦ ¹Tropical Fruit-Eating Birds and Their Food Plants: A Survey of a Costa Rican Lower Montane Forest, Nathaniel T. Wheelwright, William A. Haber, K. Greg Murray, Carlos Guindon, Biotropica Vol. 16, No. 3 (Sep., 1984), pp. 173-192
♦ ²Frugivores at a fruiting Ficus in south-eastern Peru, Jose G. Tello, Journal of Tropical Ecology (2003) 19:717–721.