Attributes / relations provided by
♦ ¹Tropical Fruit-Eating Birds and Their Food Plants: A Survey of a Costa Rican Lower Montane Forest, Nathaniel T. Wheelwright, William A. Haber, K. Greg Murray, Carlos Guindon, Biotropica Vol. 16, No. 3 (Sep., 1984), pp. 173-192
♦ ²Frugivorous Birds, Habitat Preference and Seed Dispersal in a Fragmented Costa Rican Montane Oak Forest Landscape, J.J.A.M. Wilms and M. Kappelle, Ecological Studies Vol 185, pp. 309-324
♦ ³"Fig-eating by vertebrate frugivores: a global review", MIKE SHANAHAN, SAMSON SO, STEPHEN G. COMPTON and RICHARD CORLETT, Biol. Rev. (2001), 76, pp. 529–572