Attributes / relations provided by
♦ ¹"Fig-eating by vertebrate frugivores: a global review", MIKE SHANAHAN, SAMSON SO, STEPHEN G. COMPTON and RICHARD CORLETT, Biol. Rev. (2001), 76, pp. 529–572
♦ ²del Hoyo, J., Elliott, A., Sargatal, J., Christie, D.A. & de Juana, E. (eds.). Handbook of the Birds of the World Alive. Lynx Edicions, Barcelona.
♦ ³Pteropus tonganus, Carrie A. Miller and Don E. Wilson, MAMMALIAN SPECIES No. 552, pp. 1-6 (1997)