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♦ ¹Food niche overlap among neotropical frugivorous bats in Costa Rica, Jorge E. Lopez & Christopher Vaughan, Rev. Biol. Trop. (Int. J. Trop. Biol. ISSN-0034-7744) Vol. 55 (1): 301-313, March 2007
♦ ²Dispersal of Melastome Seeds by Fruit-eating Birds of Tropical Forest Understory, Bette A. Loiselle and John G. Blake, Ecology, 80(1), 1999, pp. 330-336
♦ ³Habitat-Dependent Fruiting Behaviour of an Understorey Tree, Miconia centrodesma, and Tropical Treefall Gaps as Keystone Habitats for Frugivores in Costa Rica, Douglas J. Levey, Journal of Tropical Ecology, Vol. 6, No. 4. (Nov., 1990), pp. 409-420