Attributes / relations provided by
♦ ¹del Hoyo, J., Elliott, A., Sargatal, J., Christie, D.A. & de Juana, E. (eds.). Handbook of the Birds of the World Alive. Lynx Edicions, Barcelona.
♦ ²Patterns of movement and seed dispersal of a tropical frugivore, David A. Westcott · Devon L. Graham, Oecologia (2000) 122:249–257