Attributes / relations provided by
♦ ¹del Hoyo, J., Elliott, A., Sargatal, J., Christie, D.A. & de Juana, E. (eds.). Handbook of the Birds of the World Alive. Lynx Edicions, Barcelona.
♦ ²Tropical Fruit-Eating Birds and Their Food Plants: A Survey of a Costa Rican Lower Montane Forest, Nathaniel T. Wheelwright, William A. Haber, K. Greg Murray, Carlos Guindon, Biotropica Vol. 16, No. 3 (Sep., 1984), pp. 173-192