World Species : Protected Area : Réserve Naturelle Intégrale d'Andohahela

Réserve Naturelle Intégrale d'Andohahela

Location:	Madagascar
IUCN Category:	II
Area:	180,522 acres (73,055 hectares)
Species:	All Endangered Invasive
Website:	View website
Climate:	View Climate Data
Land Use:	View Land Use Data
Köppen climate classification:	Cfa (Climate: warm temperate; Precipitation: fully humid; Temperature: hot summer)

Ecoregions

Madagascar lowland forests
Madagascar spiny thickets
Madagascar subhumid forests

Soils

— Protected Area Boundary

	A					B				C

Share	40 %	20 %	20 %	10 %	10 %	50 %	20 %	20 %	10 %	50 %	20 %	20 %	10 %
Soil Type	Chromic Cambisols	Ferric Luvisols	Chromic Luvisols	Luvic Kastanozems	Haplic Kastanozems	Orthic Ferralsols	Humic Ferralsols	Humic Acrisols	Humic Gleysols	Dystric Regosols	Lithosols	Ferralic Cambisols	Rankers
Topsoil Texture	Medium	Medium	Medium	Medium	Medium	Medium	Medium	Medium	Medium	Medium	Medium	Medium	Medium
Reference Depth	100 cm	100 cm	100 cm	100 cm	100 cm	100 cm	100 cm	100 cm	100 cm	100 cm	10 cm	100 cm	30 cm
Drainage	Moderately well	Moderately well	Moderately well	Moderately well	Moderately well	Moderately well	Moderately well	Moderately well	Poor	Moderately well	Imperfectly	Moderately well	Imperfectly
Available Water Capacity	150 m	150 m	150 m	150 m	150 m	150 m	150 m	150 m	150 m	150 m	15 mm	150 m	50 mm
Topsoil Properties
Gravel	10 %	11 %	9 %	8 %	8 %	3 %	3 %	5 %	4 %	20 %	26 %	1 %	10 %
Sand	48 %	56 %	47 %	36 %	34 %	60 %	53 %	46 %	33 %	42 %	43 %	52 %	48 %
Silt	30 %	20 %	29 %	41 %	45 %	14 %	20 %	30 %	45 %	37 %	34 %	25 %	31 %
Clay	22 %	24 %	24 %	23 %	21 %	26 %	27 %	24 %	22 %	21 %	23 %	23 %	21 %
USDA Texture	Loam	Sandy clay loam	Loam	Loam	Loam	Sandy clay loam	Sandy clay loam	Loam	Loam	Loam	Loam	Sandy clay loam	Loam
Reference Bulk Density	1.41 g/cm³	1.41 g/cm³	1.39 g/cm³	1.38 g/cm³	1.39 g/cm³	1.41 g/cm³	1.39 g/cm³	1.39 g/cm³	1.38 g/cm³	1.4 g/cm³	1.39 g/cm³	1.41 g/cm³	1.42 g/cm³
Bulk Density	1.3 g/cm³	1.48 g/cm³	1.54 g/cm³	1.2 g/cm³	1.29 g/cm³	1.4 g/cm³	1.2 g/cm³	1.3 g/cm³	1.2 g/cm³	1.37 g/cm³	1.3 g/cm³	1.31 g/cm³	1.45 g/cm³
Organic Carbon	0.98 %	0.74 %	0.86 %	1.27 %	1.28 %	1.06 %	2.34 %	1.73 %	3.71 %	1.37 %	1.4 %	0.95 %	1.43 %
PH	6.6	6.3	6.4	7.3	7.4	5	4.8	5	5.5	5.1	7.6	4.7	5.1
Clay Cation Exchange Capacity	38 cmol kg⁻¹	23 cmol kg⁻¹	38 cmol kg⁻¹	66 cmol kg⁻¹	81 cmol kg⁻¹	8 cmol kg⁻¹	8 cmol kg⁻¹	19 cmol kg⁻¹	33 cmol kg⁻¹	23 cmol kg⁻¹	42 cmol kg⁻¹	16 cmol kg⁻¹	18 cmol kg⁻¹
Soil Cation Exchange Capacity	12 cmol kg⁻¹	7 cmol kg⁻¹	12 cmol kg⁻¹	23 cmol kg⁻¹	21 cmol kg⁻¹	6 cmol kg⁻¹	8 cmol kg⁻¹	9 cmol kg⁻¹	18 cmol kg⁻¹	11 cmol kg⁻¹	15 cmol kg⁻¹	6 cmol kg⁻¹	9 cmol kg⁻¹
Base Saturation	84 %	85 %	89 %	100 %	100 %	38 %	20 %	27 %	34 %	22 %	100 %	39 %	47 %
Total Exchangeable Bases	10.5 cmol kg⁻¹	6.1 cmol kg⁻¹	10 cmol kg⁻¹	18.4 cmol kg⁻¹	18.4 cmol kg⁻¹	2 cmol kg⁻¹	1.2 cmol kg⁻¹	2.5 cmol kg⁻¹	3.1 cmol kg⁻¹	2.6 cmol kg⁻¹	15.6 cmol kg⁻¹	2.6 cmol kg⁻¹	3.7 cmol kg⁻¹
Calcium Carbonate	0.1 %	0 %	0 %	4.5 %	5.4 %	0 %	0 %	0 %	0 %	0 %	3.9 %	0 %	0 %
Calcium Sulphate	0 %	0 %	0 %	0 %	0 %	0 %	0 %	0 %	0 %	0 %	0 %	0 %	0 %
Exchangeable Sodium	2 %	2 %	1 %	3 %	1 %	3 %	2 %	1 %	2 %	2 %	4 %	2 %	2 %
Electrical Conductivity	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	1.6 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1
Subsoil Properties
Gravel	9 %	10 %	8 %	5 %	5 %	3 %	3 %	7 %	5 %	35 %		5 %
Sand	46 %	44 %	39 %	29 %	36 %	51 %	53 %	41 %	37 %	44 %		48 %
Silt	27 %	19 %	27 %	42 %	43 %	14 %	18 %	26 %	35 %	35 %		24 %
Clay	27 %	37 %	34 %	29 %	21 %	35 %	29 %	33 %	28 %	21 %		28 %
USDA Texture	Sandy clay loam	Clay loam	Clay loam	Clay loam	Loam	Sandy clay	Sandy clay loam	Clay loam	Clay loam	Loam		Sandy clay loam
Reference Bulk Density	1.37 g/cm³	1.32 g/cm³	1.33 g/cm³	1.33 g/cm³	1.39 g/cm³	1.35 g/cm³	1.38 g/cm³	1.34 g/cm³	1.35 g/cm³	1.41 g/cm³		1.37 g/cm³
Bulk Density	1.4 g/cm³	1.47 g/cm³	1.52 g/cm³	1.18 g/cm³	1.42 g/cm³	1.37 g/cm³	1.17 g/cm³	1.35 g/cm³	1.46 g/cm³	1.53 g/cm³		1.3 g/cm³
Organic Carbon	0.4 %	0.34 %	0.38 %	0.57 %	0.51 %	0.35 %	1.12 %	0.77 %	0.69 %	0.6 %		0.35 %
PH	6.8	6	6.3	7.9	7.9	5	5	5.2	5.6	4.9		4.9
Clay Cation Exchange Capacity	41 cmol kg⁻¹	21 cmol kg⁻¹	40 cmol kg⁻¹	68 cmol kg⁻¹	97 cmol kg⁻¹	9 cmol kg⁻¹	7 cmol kg⁻¹	15 cmol kg⁻¹	41 cmol kg⁻¹	18 cmol kg⁻¹		12 cmol kg⁻¹
Soil Cation Exchange Capacity	14 cmol kg⁻¹	8 cmol kg⁻¹	15 cmol kg⁻¹	24 cmol kg⁻¹	19 cmol kg⁻¹	4 cmol kg⁻¹	5 cmol kg⁻¹	7 cmol kg⁻¹	11 cmol kg⁻¹	6 cmol kg⁻¹		5 cmol kg⁻¹
Base Saturation	95 %	77 %	93 %	100 %	100 %	46 %	40 %	21 %	62 %	25 %		57 %
Total Exchangeable Bases	11.1 cmol kg⁻¹	6.6 cmol kg⁻¹	12.3 cmol kg⁻¹	21.1 cmol kg⁻¹	19.1 cmol kg⁻¹	2.8 cmol kg⁻¹	4.1 cmol kg⁻¹	3 cmol kg⁻¹	9 cmol kg⁻¹	2.1 cmol kg⁻¹		3.7 cmol kg⁻¹
Calcium Carbonate	0.1 %	0 %	0 %	10 %	10.4 %	0 %	0 %	0 %	0 %	0 %		0 %
Calcium Sulphate	0 %	0 %	0 %	0 %	0 %	0 %	0 %	0 %	0 %	0 %		0 %
Exchangeable Sodium	1 %	2 %	1 %	3 %	3 %	3 %	3 %	1 %	4 %	2 %		2 %
Electrical Conductivity	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	3.6 dS m-1	2.2 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1	0.1 dS m-1		0.1 dS m-1

Soil data provided by FAO/IIASA/ISRIC/ISSCAS/JRC, 2012. Harmonized World Soil Database (version 1.2). FAO, Rome, Italy and IIASA, Laxenburg, Austria.

Köppen climate classification provided by Kottek, M., J. Grieser, C. Beck, B. Rudolf, and F. Rubel, 2006: World Map of Köppen-Geiger Climate Classification updated. Meteorol. Z., 15, 259-263