Peronosporaceae are a family of water moulds that contains 17 genera, comprising more than 600 species. Most of them are called downy mildews. The closest relative of Peronosporaceae is Phytophthora. Peronosporaceae are obligate biotrophic plant pathogens. They parasitize their host plants as an intercellular mycelium using haustoria to penetrate the host cells. The downy mildews reproduce asexually by forming sporangia on distinctive white sporangiophores usually formed on the lower surface of infected leaves. These constitute the "downy mildew". The sporangia are wind-dispersed to the surface of other leaves. According to the genus concerned, the sporangia may then germinate by forming zoospores, thus resembling Phytophthora, or by germ-tube. In the latter case, the sporangia behave as c