Attributes / relations provided by
♦ ¹Tropical Fruit-Eating Birds and Their Food Plants: A Survey of a Costa Rican Lower Montane Forest, Nathaniel T. Wheelwright, William A. Haber, K. Greg Murray, Carlos Guindon, Biotropica Vol. 16, No. 3 (Sep., 1984), pp. 173-192
♦ ²Food niche overlap among neotropical frugivorous bats in Costa Rica, Jorge E. Lopez & Christopher Vaughan, Rev. Biol. Trop. (Int. J. Trop. Biol. ISSN-0034-7744) Vol. 55 (1): 301-313, March 2007
♦ ³Habitat-Dependent Fruiting Behaviour of an Understorey Tree, Miconia centrodesma, and Tropical Treefall Gaps as Keystone Habitats for Frugivores in Costa Rica, Douglas J. Levey, Journal of Tropical Ecology, Vol. 6, No. 4. (Nov., 1990), pp. 409-420